Squamulose terricolous on basic and siliceous soils.
Nota: Perithecia containing cells of the thalline algae. Asci bi-spored. Muriform spores pale to dark-brown, broadly ellipsoidal, elongate ellipsoidal or subcilindrical, 29-50 x 14-28 micras:
Undersurface dark brown, anchored by concolorous branched thick rhizines:
Squamules matt, pale, with perithecia immersed, 1-10 per squamule:
Altitude: 860 m